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Ces règles concernent les candidatures, je les ai toutes lues.

Ces règles concernent les candidatures, je les ai toutes lues.
These rules concern the applications, I read all of them (f).
From Lesson 436:The pronouns all and everyone (tous, toutes)·French

Questions & Answers about Ces règles concernent les candidatures, je les ai toutes lues.

Why is it toutes instead of tout, and what exactly does it refer to?
toutes is a feminine plural pronoun meaning "all of them." It refers back to les candidatures (the applications), which is feminine plural. If it referred to a masculine plural noun, you would use tous.
Where exactly does toutes have to go in this sentence?
When toutes (or tous) is paired with a direct object pronoun like les in a compound tense like the passé composé, it must go between the helping verb (ai) and the past participle (lues). You cannot put it at the very end of the sentence.
The lesson says we pronounce the final "s" for the pronoun tous. Do we pronounce the "s" in toutes here?
No, the final "s" in toutes is always silent. You just pronounce the final "t" sound. The special rule about pronouncing the final "s" only applies to the masculine plural pronoun tous.
Why does the past participle lues have an -es at the end?
This is the preceding direct object rule. In the passé composé, when a direct object comes before the verb, the past participle must agree with it in gender and number. Here, the direct object pronoun les (replacing the feminine plural candidatures) comes before ai, so lu gets an -es to become lues.

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