Chinese, Mandarin

Spanish

English

Hindi

Arabic

Portuguese

Bengali

Russian

Japanese

Punjabi

German

Javanese

Malay

Telugu

Vietnamese

Korean

French

Marathi

Tamil

Urdu

Turkish

Italian

Chinese, Yue / Cantonese

Thai

Gujarati

Chinese, Jin

Chinese, Min Nan

Persian, Dari

Persian, Western

Polish

Pashto

Kannada

Chinese, Xiang

Malayalam

Sundanese

Hausa

Odia

Burmese

Chinese, Hakka

Ukrainian

Bihari, Bhojpuri

Tagalog

Yoruba

Bihari, Maithili

Uzbek

Sindhi

Amharic

Fula

Romanian

Oromo

Igbo

Azerbaijani

Chinese, Gan

Dutch

Kurdish, Central

Kurdish, Northern

Kurdish, Southern

Malagasy

Lahnda, Saraiki

Nepali

Sinhalese

Chittagonian

Zhuang

Khmer

Turkmen

Assamese

Madurese

Somali

Marwari

Bihari, Magahi

Haryanvi

Hungarian

Chhattisgarhi

Greek, Modern

Chewa

Dakhini

Akan, Twi

Akan, Fante

Kazakh

Chinese, Min Bei

Sylheti

Zulu

Czech

Kinyarwanda

Marwari, Dhundhari

Haitian Creole

Chinese, Min Dong

Ilocano

Quechua

Kirundi

Swedish

Hmong

Shona

Uyghur

Hiligaynon

Mossi

Xhosa

Belarusian

Balochi

Anthropology

<OpenstaxParagraph>
	The Denisovans, like <OpenstaxEm>Homo naledi</OpenstaxEm>, are archaic <OpenstaxEm>Homo</OpenstaxEm>. There are not a lot of specimens—just one finger bone, three teeth, some <a href="https://openstax.org/r/Long_bone">long bone</a> fragments, a partial jawbone, and a <a href="https://openstax.org/r/Parietal_bone">parietal bone</a> skull fragment. Because of this lack of evidence, very little is known of their anatomical features. Some of the specimens come from <OpenstaxTerm>Denisova Cave</OpenstaxTerm> in Siberia, Russia, and are dated to between 500,000 and 30,000 years ago. These dates are arrived at based on the few fossils that exist, inferences made from genetic studies, and sediment analysis. More recently another specimen was found on the Tibetan plateau. In 1980 a jaw and two teeth were uncovered in the Baishiya Karst Cave by a monk, but it wasn’t until 2010 that scientists were able to study the jaw. Dating placed the specimen at approximately 160,000 years ago. Protein analysis determined the jaw to be of Denisovan origin and from a member of a population who were most likely well adapted to living in high altitudes (Chen et al. 2019).
</OpenstaxParagraph>
<OpenstaxParagraph>
	Because so few bones have been found, most understanding of this species comes from genetic analyses. According to nuclear DNA studies, Denisovans and Neanderthals were more closely related to each other than they were to modern humans. DNA evidence suggests that the Denisovans interbred with modern humans and with local Neanderthal populations over multiple time periods. Tracing the male Y chromosome, one study indicated that interbreeding between early humans and Neanderthals actually replaced the ancient Denisovan Y chromosome once found in Neanderthals. The time of divergence of the Denisovan is estimated to be around 700,000 years ago, with modern humans diverging from the Neanderthal around 370,000 years ago (Petr et al. 2020). <OpenstaxEm>H. heidelbergensis</OpenstaxEm> is typically considered to have been the direct ancestor of both Denisovans and Neanderthals, and sometimes also of modern humans.
</OpenstaxParagraph>
<OpenstaxParagraph>
	One specimen is a first-generation hybrid, <OpenstaxTerm>Denisova 11</OpenstaxTerm>—nicknamed “<a href="https://openstax.org/r/Denny_hybrid_hominin">Denny</a>”—that had a Denisovan father and a Neanderthal mother (Slon et al<OpenstaxEm>.</OpenstaxEm> 2018). Denisova 11 was found in Denisova Cave in Russia and provides evidence that Late Pleistocene <OpenstaxEm>Homo</OpenstaxEm> species interbred when the groups met. Comparison of the DNA of these three groups suggest that most modern-day Europeans and Asians inherited about 1–4 percent of their DNA from Neanderthals, with no Denisovan ancestry in Europe and 0.1 percent in China. The genetics found in Tibetans, Melanesians, and Indigenous Australian are currently being challenged; originally, they were thought to be about 3–5 percent Denisovan and 2.74 percent Neanderthal. Statistical geneticist Ryan <OpenstaxTerm>Bohlender</OpenstaxTerm> and his team have investigated the percentages of extinct hominin DNA in modern humans. They concluded that Neanderthals and Denisovans are not the whole story and that there could be a third group yet unknown contributing to the Pacific Islander <OpenstaxTerm>genome</OpenstaxTerm> (Rogers, Bohlender, and Huff 2017). Statistical and genetic evidence can serve as indicators of the existence of a group for which no fossils have yet been found. These are referred to as <OpenstaxTerm>ghost populations</OpenstaxTerm>. For example, there are indications that 2–19 percent of the DNA of four West African populations may have come from an unknown archaic hominin that split from the ancestor of humans and Neanderthals between 360 KYA and 1.02 MYA (Durvasula and Sankararaman 2020). The hypothesis of a third lineage in the genus <OpenstaxEm>Homo</OpenstaxEm> appears to have received further confirmation with a discovery in China.
</OpenstaxParagraph>

5.3.5 The Denisovans

1 What Is Anthropology?

1.1 The Study of Humanity, or "Anthropology Is Vast"

1.1.1 The Heart of Anthropology: Central Narrative and Commitments

1.1.2 Central Commitment #1: Exploring Sociocultural Diversity

1.1.3 Central Commitment #2: Understanding How Societies Hold Together

1.1.4 Central Commitment #3: Examining the Interdependence of Humans and Nature

1.2 The Four-Field Approach: Four Approaches within the Guiding Narrative

1.2.1 Biological Anthropology

1.2.2 Archaeology

1.2.3 Cultural Anthropology

1.2.4 Linguistic Anthropology

1.2.5 How the Four Fields Work Together: The Example of Race

1.3 Overcoming Ethnocentrism

1.3.1 Enculturation and Ethnocentrism

1.3.2 Primitivism and Orientalism

1.4 Western Bias in Our Assumptions about Humanity

1.4.1 Primitivism and Orientalism in Popular Culture

1.4.2 The Bias of Backwardness

1.5 Holism, Anthropology’s Distinctive Approach

1.5.1 Society as an Integrated Whole

1.5.2 Sources of Contradiction, Conflict, and Change

1.6 Cross-Cultural Comparison and Cultural Relativism

1.6.1 Relativism Is Not “Anything Goes”

1.6.2 Morality, Activism, and Cultural Relativism

1.7 Reaching for an Insider’s Point of View

1.7.1 The Challenge of Representing Others

1.7.2 Collaborative Methods of Representation

1.7.3 Working across Cultures toward Common Goals

1.7.4 Suggested Readings

2 Methods: Cultural and Archaeological

2.1 Archaeological Research Methods

2.1.1 Archaeological Techniques

2.1.2 Archaeological Dating Methods

2.2 Conservation and Naturalism

2.2.1 Early Efforts

2.2.2 Salvage Anthropology

2.2.3 Museum Collections

2.2.4 Interpretation and Voice

2.3 Ethnography and Ethnology

2.3.1 The Development of Ethnography and Ethnology

2.3.2 Perspective and Interpretation in Ethnography

2.4 Participant Observation and Interviewing

2.4.1 Participant Observation

2.4.2 Interviewing Informants

2.4.3 Ethical Considerations

2.5 Quantitative and Qualitative Analysis

2.5.1 Differences between Quantitative and Qualitative Information

2.5.2 Modeling

2.5.3 The Science of Anthropology

2.6 Collections

2.6.1 Archives

2.6.2 Three-Dimensional Collections

2.6.3 Ownership

2.6.4 Suggested Readings

3 Culture Concept Theory: Theories of Cultural Change

3.1 The Homeyness of Culture

3.2 The Winkiness of Culture

3.3 The Elements of Culture

3.3.1 Culture Is What We Make

3.3.2 Culture Is What We Do

3.3.3 Culture Is What We Think

3.4 The Aggregates of Culture

3.4.1 Symbols

3.4.2 Ritual

3.4.3 Social Structure

3.5 Modes of Cultural Analysis

3.5.1 Evolution, Adaptation, and Historical Particularism

3.5.2 Functionalism

3.5.3 Structuralism

3.5.4 Ontology

3.6 The Paradoxes of Culture

3.6.1 Paradox 1: Culture Is Continuous, but It Changes

3.6.2 Paradox 2: Culture Is Bounded but Mobile

3.6.3 Paradox 3: Culture Is Consensual but Contested

3.6.4 Paradox 4: Culture Is Shared, but It Varies

3.6.5 Suggested Readings

4 Biological Evolution and Early Human Evidence

4.1 What Is Biological Anthropology?

4.1.1 Looking to the Deep Past

4.1.2 Exploring What It Means to Be Human

4.2 What’s in a Name? The Science of Taxonomy

4.2.1 Defining the Science of Taxonomy

4.2.2 Defining a Species

4.3 It’s All in the Genes! The Foundation of Evolution

4.3.1 The Units of Life

4.3.2 Gregor Mendel and the Laws of Heredity

4.3.3 Mendelian Inheritance in Humans

4.4 Evolution in Action: Past and Present

4.4.1 Early Evolutionists and the Fixity of Species

4.4.2 Charles Darwin’s Role in Changing Views of the Natural World

4.4.3 Understanding Darwin’s Theory of Natural Selection

4.5 What Is a Primate?

4.5.1 What Is a Primate?

4.5.2 Primate Behavioral Variation

4.5.3 Explaining Primate Success

4.5.4 Primate Classification and Taxonomy

4.6 Origin of and Classification of Primates

4.6.1 Understanding Concepts of Time

4.6.2 Fossils and Dating Methods